The Pantanal is one of the world's largest tropical wetland areas, and is located mostly within the Brazilian state of Mato Grosso do Sul, but it extends into Mato Grosso and portions of Bolivia and Paraguay. It sprawls over an area estimated at between 140,000 and 195,000 square kilometres (54,000 and 75,000 sq mi). The Pantanal ecosystem is also thought to be home to 1000 bird species, 400 fish species, 300 mammalian species, 480 reptile species and over 9000 different subspecies of invertebrates. Among the rarest animals to inhabit the wetland of the Pantanal are the marsh deer (Blastocerus dichotomus) and the giant river otter (Pteroneura brasiliensis). Parts of the Pantanal are also home to the following endangered or threatened species: the hyacinth macaw (Anodorhyncus hyacinthinus) (a bird endangered due to smuggling), the crowned solitary eagle, the maned wolf (Chrysocyon brachyurus), the bush dog (Speothos venaticus), the South American tapir (Tapirus terrestris), the giant anteater (Myrmecophaga tridactyla) and the yacare caiman (Caiman yacare).